红腺悬钩子Rubus sumatranus
中文名（Chinese Name）：红腺悬钩子
学名（Scientific Name）：Rubus sumatranus Miq.
英文名（English Common Name）：
别名（Chinese Common Name）：马泡、红刺苔
异名（Synonym）：Rubus dolichocephalus Hayata Rubus asper var. myriadenus (H. Lév. et Vaniot) Focke Rubus asper var. pekanius Focke Rubus indotibetanus Koidz. Rubus rosifolius subsp. sumatranus (Miq.) Focke Rubus asper subvar var. grandifoliolatus (H. Léveillé) Focke Rubus myriadenus H. Lév. et Vaniot Rubus sorbifolius Maxim. Rubus takasagoensis Koidz. Rubus myriadenus var. grandifoliolatus H. Lév. Rubus somae Hayata Rubus asper subvar. grandifoliolatus (H. Léveillé) Focke Rubus asper subvar grandifoliolatus (H.L‚v.) Focke
科属（Family & Genus）：蔷薇科（Rosaceae）悬钩子属
形态特征（Description）：直立或攀援灌木；小枝、叶轴、叶柄、花梗和花序均被紫红色腺毛、柔毛和皮刺；腺毛长短不等，长者达4-5毫米，短者1-2毫米。小叶5-7枚，稀3枚，卵状披针形至披针形， 长3-8厘米，宽1.5-3厘米，顶端渐尖，基部圆形，两面疏生柔毛，沿中脉较密，下面沿中脉有小皮刺，边缘具不整齐的尖锐锯齿；叶柄长3-5厘米，顶生小叶柄长达1厘米；托叶披针形或线状披针形，有柔毛和腺毛。花3朵或数朵成伞房状花序，稀单生；花梗长2-3厘米；苞片披针形；花直径1-2厘米；花萼被长短不等的腺毛和柔毛；萼片披针形，长0.7-1厘米，宽0.2-0.4厘米，顶端长尾尖，在果期反折；花瓣长倒卵形或匙状，白色，基部具爪；花丝线形；雌蕊数可达400，花柱和子房均无毛。果实长圆形，长1.2-1.8厘米，桔红色，无毛。花期4-6月，果期7-8月。
分布（Distribution）：产湖北、湖南、江西、安徽、浙江、福建、台湾、广东、广西、四川、贵州、云南、西藏。生山地、山谷疏密林内、林缘、灌丛内、竹林下及草丛中，海拔达2000米。朝鲜、日本、锡金、尼泊尔、印度、越南、泰国、老挝、柬埔寨、印度尼西亚也有分布。
用途（Use）：根入药，有清热、解毒、利尿之效。
引自中国植物志英文版： FOC Vol. 9 Page 225
Rubus sumatranus Miquel, Fl. Ned. India, Eerste Bijv. 307. 1860.
红腺悬钩子 hong xiao xuan gou zi| Rosaceae | Rubus
Rubus asper Wallich ex G. Don var. myriadenus (H. Léveillé & Vaniot) Focke; R. asper var. myriadenus (H. Léveillé & Vaniot) Focke subvar. grandifoliolatus (H. Léveillé) Focke; R. asper var. pekanius Focke; R. dolichocephalus Hayata; R. indotibetanus Koidzumi; R. myriadenus H. Léveillé & Vaniot; R. myriadenus var. grandifoliolatus H. Léveillé; R. rosifolius Smith subsp. sumatranus (Miquel) Focke; R. somae Hayata; R. sorbifolius Maximowicz; R. takasagoensis Koidzumi.
Shrubs erect or scandent. Branchlets brownish to dark reddish brown, cylindric, with long soft hairs, usually with scattered setose purplish red glandular hairs and curved prickles; glandular hairs and prickles unequal in length, glandular hairs to 4–5 mm, prickles to 8 mm. Leaves imparipinnate, to 15 cm, 5–7-foliolate, rarely 3-foliolate; petiole 3–5 cm, petiolule of terminal leaflet to 1 cm, lateral leaflets shortly petiolulate, petiolule and rachis soft hairy, with intermixed glandular hairs, with curved minute prickles; stipules lanceolate or linear-lanceolate, 6–8 mm, with soft hairs and intermixed glandular hairs; blade of leaflets ovate-lanceolate to lanceolate, 3–8 × 1.5–3 cm, abaxially villous, with gland-tipped hairs and small prickles along midvein, adaxially villous especially along midvein, base rounded, margin irregularly sharply serrate to doubly serrate, sometimes 3-lobed on terminal leaflet, apex acuminate. Inflorescences terminal, corymbose, 4–7 cm, 3- to several flowered, rarely flowers solitary; rachis and pedicels villous, with intermixed glandular hairs, with minute prickles; bracts lanceolate or linear, 5–7 mm, with soft hairs and intermixed glandular hairs. Pedicel 2–3 cm. Flowers 1–2 cm in diam. Calyx abaxially with soft hairs, with intermixed unequal long gland-tipped hairs; sepals reflexed in fruit, lanceolate or oblong-lanceolate, 7–12 × 2–4 mm; apex long caudate. Petals white, narrowly obovate or spatulate, slightly shorter than sepals, base clawed. Stamens many, shorter than petals. Pistils to 400; style and ovary glabrous; torus raised, oblong, base shortly stalked. Aggregate fruit orange-red, oblong, 1.2–1.8 cm × 7–11 mm, glabrous; pyrenes reticulate. Fl. Apr–Jun, fr. Jul–Aug. 2n = 14.
Forests, forest margins, thickets, bamboo forests, grasslands; 700--2500 m. Anhui, Fujian, Guangdong, Guangxi, Guizhou, Hainan, Hubei, Hunan, Jiangxi, Sichuan, Taiwan, Xizang, Yunnan, Zhejiang [Bhutan, Cambodia, N India, Indonesia, Japan, Korea, Laos, Malaysia, Myanmar, Nepal, Sikkim, Thailand, Vietnam].
（责任编辑：徐晔春）
学名（Scientific Name）：Rubus sumatranus Miq.
英文名（English Common Name）：
别名（Chinese Common Name）：马泡、红刺苔
异名（Synonym）：Rubus dolichocephalus Hayata Rubus asper var. myriadenus (H. Lév. et Vaniot) Focke Rubus asper var. pekanius Focke Rubus indotibetanus Koidz. Rubus rosifolius subsp. sumatranus (Miq.) Focke Rubus asper subvar var. grandifoliolatus (H. Léveillé) Focke Rubus myriadenus H. Lév. et Vaniot Rubus sorbifolius Maxim. Rubus takasagoensis Koidz. Rubus myriadenus var. grandifoliolatus H. Lév. Rubus somae Hayata Rubus asper subvar. grandifoliolatus (H. Léveillé) Focke Rubus asper subvar grandifoliolatus (H.L‚v.) Focke
科属（Family & Genus）：蔷薇科（Rosaceae）悬钩子属
形态特征（Description）：直立或攀援灌木；小枝、叶轴、叶柄、花梗和花序均被紫红色腺毛、柔毛和皮刺；腺毛长短不等，长者达4-5毫米，短者1-2毫米。小叶5-7枚，稀3枚，卵状披针形至披针形， 长3-8厘米，宽1.5-3厘米，顶端渐尖，基部圆形，两面疏生柔毛，沿中脉较密，下面沿中脉有小皮刺，边缘具不整齐的尖锐锯齿；叶柄长3-5厘米，顶生小叶柄长达1厘米；托叶披针形或线状披针形，有柔毛和腺毛。花3朵或数朵成伞房状花序，稀单生；花梗长2-3厘米；苞片披针形；花直径1-2厘米；花萼被长短不等的腺毛和柔毛；萼片披针形，长0.7-1厘米，宽0.2-0.4厘米，顶端长尾尖，在果期反折；花瓣长倒卵形或匙状，白色，基部具爪；花丝线形；雌蕊数可达400，花柱和子房均无毛。果实长圆形，长1.2-1.8厘米，桔红色，无毛。花期4-6月，果期7-8月。
分布（Distribution）：产湖北、湖南、江西、安徽、浙江、福建、台湾、广东、广西、四川、贵州、云南、西藏。生山地、山谷疏密林内、林缘、灌丛内、竹林下及草丛中，海拔达2000米。朝鲜、日本、锡金、尼泊尔、印度、越南、泰国、老挝、柬埔寨、印度尼西亚也有分布。
用途（Use）：根入药，有清热、解毒、利尿之效。
引自中国植物志英文版： FOC Vol. 9 Page 225
Rubus sumatranus Miquel, Fl. Ned. India, Eerste Bijv. 307. 1860.
红腺悬钩子 hong xiao xuan gou zi| Rosaceae | Rubus
Rubus asper Wallich ex G. Don var. myriadenus (H. Léveillé & Vaniot) Focke; R. asper var. myriadenus (H. Léveillé & Vaniot) Focke subvar. grandifoliolatus (H. Léveillé) Focke; R. asper var. pekanius Focke; R. dolichocephalus Hayata; R. indotibetanus Koidzumi; R. myriadenus H. Léveillé & Vaniot; R. myriadenus var. grandifoliolatus H. Léveillé; R. rosifolius Smith subsp. sumatranus (Miquel) Focke; R. somae Hayata; R. sorbifolius Maximowicz; R. takasagoensis Koidzumi.
Shrubs erect or scandent. Branchlets brownish to dark reddish brown, cylindric, with long soft hairs, usually with scattered setose purplish red glandular hairs and curved prickles; glandular hairs and prickles unequal in length, glandular hairs to 4–5 mm, prickles to 8 mm. Leaves imparipinnate, to 15 cm, 5–7-foliolate, rarely 3-foliolate; petiole 3–5 cm, petiolule of terminal leaflet to 1 cm, lateral leaflets shortly petiolulate, petiolule and rachis soft hairy, with intermixed glandular hairs, with curved minute prickles; stipules lanceolate or linear-lanceolate, 6–8 mm, with soft hairs and intermixed glandular hairs; blade of leaflets ovate-lanceolate to lanceolate, 3–8 × 1.5–3 cm, abaxially villous, with gland-tipped hairs and small prickles along midvein, adaxially villous especially along midvein, base rounded, margin irregularly sharply serrate to doubly serrate, sometimes 3-lobed on terminal leaflet, apex acuminate. Inflorescences terminal, corymbose, 4–7 cm, 3- to several flowered, rarely flowers solitary; rachis and pedicels villous, with intermixed glandular hairs, with minute prickles; bracts lanceolate or linear, 5–7 mm, with soft hairs and intermixed glandular hairs. Pedicel 2–3 cm. Flowers 1–2 cm in diam. Calyx abaxially with soft hairs, with intermixed unequal long gland-tipped hairs; sepals reflexed in fruit, lanceolate or oblong-lanceolate, 7–12 × 2–4 mm; apex long caudate. Petals white, narrowly obovate or spatulate, slightly shorter than sepals, base clawed. Stamens many, shorter than petals. Pistils to 400; style and ovary glabrous; torus raised, oblong, base shortly stalked. Aggregate fruit orange-red, oblong, 1.2–1.8 cm × 7–11 mm, glabrous; pyrenes reticulate. Fl. Apr–Jun, fr. Jul–Aug. 2n = 14.
Forests, forest margins, thickets, bamboo forests, grasslands; 700--2500 m. Anhui, Fujian, Guangdong, Guangxi, Guizhou, Hainan, Hubei, Hunan, Jiangxi, Sichuan, Taiwan, Xizang, Yunnan, Zhejiang [Bhutan, Cambodia, N India, Indonesia, Japan, Korea, Laos, Malaysia, Myanmar, Nepal, Sikkim, Thailand, Vietnam].
（责任编辑：徐晔春）
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